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J. C. Caruthers and Debinski, D., Montane Meadow Butterfly Species Distributions in the Greater Yellowstone Ecosystem, University of Wyoming National Park Service Research Center Annual Report, vol. 30, no. 1, pp. 85-96, 2006.
J. C. Caruthers, Debinski, D. M., and Whickam, H., CHAPTER TWO: BUTTERFLY INDICATORS OF MONTANE MEADOWS ALONG A HYDROLOGICAL GRADIENT: RESPONSES FROM A SPATIAL AND TEMPORAL SCALE, Butterfly Community Temporal Trends and Responses to Resource Availability Along a Hydrologic Gradient of Montane Meadows, p. 8, 2008.
J. C. Caruthers and Debinski, D. M., CHAPTER THREE: BUTTERFLY RESPONSES TO FLORAL AND HOST PLANT RESOURCES IN MONTANE MEADOWS, Butterfly Community Temporal Trends and Responses to Resource Availability Along a Hydrologic Gradient of Montane Meadows, p. 30, 2008.